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I think a part that is missing from this is suspension design. Some designs benefit greatly from a steeper seat tube angle and others are negatively impacted. If your suspension sucks down when you climb a steeper angle helps, if your bike firms up it doesn't help.
The part about muscle memory is HUGE, I rode my wife's bike (which has a steeper STA) and found my legs were sore in odd places the next day. Oddly I also felt like her bike was a rocket ship climbing, it felt like I was creating way more power than normal. Riding flat and down hill her bike feels like complete dog poo to me. I ride a 1st gen Hightower and my wife has a 4th gen 5010. The jury is still out on if it is the geo or the smaller wheels that make me dislike the 5010 so much.
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